Spatial organization of ecologically-relevant high order flow properties and implications for river habitat assessment

The turbulent properties of flow in rivers are of fundamental importance to aquatic organisms yet are rarely quantified during routine river habitat assessment surveys or the design of restoration schemes due to their complex nature. This thesis uses a detailed review of the literature to highlight the various ways in which plants and animals modify the flow field, how this can deliver beneficial effects; and how turbulence can also generate threats to growth and survival. The thesis then presents the results from detailed field assessments of turbulence properties undertaken on low, intermediate and high gradient rivers to advance scientific understanding of the hydrodynamics of rivers and inform effective habitat assessment and restoration. A reach-scale comparison across sites reveals spatial variations in the relationships between turbulent parameters, emphasising the need for direct measurement of turbulence properties, while a geomorphic unit scale assessment suggests that variations in turbulence at the scale of individual roughness elements, and/or within the same broad groupings of geomorphic units (e.g. different types of pools) can have an important influence on hydraulic habitat. The importance of small-scale flow obstructions is further emphasised through analysis of the temporal dynamics of turbulence properties with changes in flow stage and vegetation growth. The highest magnitude temporal changes in turbulence properties were associated with individual boulders and vegetation patches respectively, indicating flow intensification around these sub-geomorphic unit scale features. Experimental research combining flow measurement with underwater videography reveals that more sophisticated turbulence parameters provide a better explanation of fish behaviour and habitat use under field conditions, further supporting direct measurement of turbulent properties where possible. The new insights into interactions between geomorphology, hydraulics and aquatic organisms generated by this work offer opportunities for refining habitat assessment and restoration design protocols to better integrate the important role of turbulence in generating suitable physical habitat for aquatic organisms.